1. Department of Zoology, University of British Columbia, Vancouver, British Columbia, Canada V6T 1Z4
2. Laboratoire de Botanique évolutive, Institut de biologie, Emile Argand 11, 2009 Neuchâtel, Switzerland
3. Swedish Board of Fisheries, Institute of Coastal Research, Skolgatan 6, 742 42 Öregrund, Sweden
4. Department of Biological Sciences, University of Calgary, Calgary, Alberta, Canada T2N 1N4