1. Key Laboratory of Plant Resource Conservation and Sustainable Utilization, The Chinese Academy of Sciences, South China Botanical Garden, Guangzhou 510650, People's Republic of China
2. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Guangzhou 510650, People's Republic of China
3. Systematic Botany and Mycology, University of Munich (LMU), 80638 Munich, Germany