1. Smithsonian Tropical Research Institute, Apartado 0843-03092, Balboa, Ancon, Republic of Panama
2. Division of Marine Biotechnology, Oceanography Section, Science Research Centre, Kochi University, 200 Monobe, Nankoku, Kochi 783-8502, Japan
3. Department of Ecology and Evolutionary Biology, University of California, Los Angeles, CA 90095, USA
4. Marine Science Institute and Department of Ecology, Evolution and Marine Biology University of California, Santa Barbara, CA 93106, USA