1. UNM Biology Department, University of New Mexico, MSC03-2020, Albuquerque, NM 87131-0001, USA
2. Department of Biology, University of Massachusetts, Amherst, MA 01003, USA
3. DST-NRF Centre of Excellence at the PercyFitz Patrick Institute, Department of Zoology and Entomology, University of Pretoria, Private Bag X20, Hatfield 0028, South Africa