1. Behavioural Ecology, Department of Biological Sciences, Macquarie University, Sydney, New South Wales 2109, Australia
2. CIBIO Research Centre in Biodiversity and Genetic Resources, InBIO, Universidade do Porto, Campus Agrário de Vairão, Rua Padre Armando Quintas, No. 7, 4485-661 Vairão, Vila do Conde, Portugal