1. School of Aquatic and Fishery Sciences, University of Washington, 1122 NE Boat Street, Seattle, WA 98105, USA
2. National Marine Fisheries Service, Northwest Fisheries Science Center, 2725 Montlake Boulevard East, Seattle, WA 98112, USA
3. School of Biology and Ecology, University of Maine, Murray Hall, Orono, ME 04469, USA