1. Université de Lyon, 69000, Lyon; Université Lyon 1; CNRS, UMR 5558, Laboratoire de Biométrie et Biologie Évolutive, 69622 Villeurbanne, France
2. Department of Ecology and Natural Resource Management, Norwegian University of Life Sciences, 1432 Aas, Norway
3. Department for Arctic Ecology, Norwegian Institute for Nature Research, Fram Centre, 9296 Tromsø, Norway
4. The James Hutton Institute, Aberdeen AB15 8QH, UK
5. Department for Terrestrial Ecology, Norwegian Institute for Nature Research, 7485 Trondheim, Norway
6. Department of Production Animal Clinical Sciences, Norwegian University of Life Sciences, 0033 Oslo, Norway