1. Field Station Fabrikschleichach, Biozentrum, University of Würzburg, Glashüttenstrasse 5, Rauhenebrach 96181, Germany
2. School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4814, Australia
3. Zoology Building, School of Biological Sciences, University of Aberdeen, Tillydrone Avenue, Aberdeen AB24 2TZ, UK