1. Centre for Evolutionary Biology, School of Animal Biology, University of Western Australia, Crawley, Western Australia 6009, Australia
2. Dipartimento di Biologia, Università di Padova, via U. Bassi 58/B, 35131 Padova, Italy
3. School of Biological Sciences, University of Auckland, Auckland 1142, New Zealand
4. Department of Life Sciences, University of the West Indies, St Augustine, Trinidad and Tobago
5. Department of Biological Sciences, University of Windsor, 401 Sunset Avenue, Windsor, Ontario, Canada N9B 3P4