1. Norwegian Institute for Nature Research, FRAM-High North Research Centre for Climate and the Environment, 9296 Tromsø, Norway
2. Centre for Biodiversity Dynamics, Norwegian University of Science and Technology (NTNU), 7491 Trondheim, Norway
3. Department of Natural Sciences, Tromsø University Museum, 9037 Tromsø, Norway
4. Norwegian Polar Institute, FRAM-High North Research Centre for Climate and the Environment, 9296 Tromsø, Norway