1. Department of Ecology and Evolutionary Biology, University of California, 621 Charles E. Young Drive South, Los Angeles, CA 90095-1606, USA
2. Department of Entomology, North Carolina State University, Campus Box 7613, Raleigh, NC 27695-7613, USA
3. Department of Biological Sciences, Dominican University, 7900 West Division St., River Forest, IL 60305, USA