1. Department of Biology, KU Leuven, Laboratory of Socioecology and Social Evolution, Leuven, Belgium
2. Department of Biology, KU Leuven, Functional Genomics and Proteomics Group, Leuven, Belgium
3. Institute for Evolution and Biodiversity, University of Münster, Molecular Evolution and Sociobiology Group, Münster, Germany
4. Departments of Entomology and Chemistry, University of California, Riverside, CA 92521, USA