1. Institut de Biologia Evolutiva (CSIC-UPF), Passeig Marítim de la Barceloneta 37, 08003 Barcelona, Catalonia, Spain
2. Laboratoire d’Écologie Alpine, Univ. Grenoble Alpes, CNRS, Univ. Savoie Mont Blanc, CNRS, LECA, Grenoble 38000, France
3. Department of Organismic and Evolutionary Biology and Museum of Comparative Zoology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA