1. MOE Key Laboratory of Biosystems Homeostasis & Protection, College of Life Sciences, Zhejiang University, Hangzhou, Zhejiang 310058, People's Republic of China
2. Zhejiang Zhoushan Archipelago Observation and Research Station, Institute of Eco-Chongming, Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, People's Republic of China
3. Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yunnan 666303, People's Republic of China
4. Section for Molecular Ecology and Evolution, Globe Institute, University of Copenhagen, Copenhagen, Denmark