1. CSIRO Environment, GPO Box 1700, Canberra, Australian Capital Territory 2601, Australia
2. Research School of Biology, Australian National University, Canberra, Australian Capital Territory 0200, Australia
3. Department of Zoology and Environmental Management, University of Kelaniya, Kelaniya 11600, Sri Lanka
4. Centre for Integrative Ecology, School of Life and Environmental Sciences, Deakin University, Burwood, Victoria 3125, Australia
5. Centre d'Etudes Biologiques de Chizé UMR 7372 Université de la Rochelle-CNRS, 405 route de Prissé la Charrière 79360 Villiers en Bois, France
6. CSIRO Environment, University of Tasmania, College Road, Sandy Bay Tas 7005, Australia