1. Department of Anthropology, University College London, 14 Taviton Street, London WC1H 0BW, UK
2. The Institute of Zoology, Zoological Society of London, Regent's Park, London NW8 7LS, UK
3. Centre for Biodiversity and Environment Research, Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, UK
4. School of Biological and Chemical Sciences, Queen Mary University of London, London E1 4NS, UK