1. Molecular Ecology and Fisheries Genetics Laboratory, Bangor University, Bangor, Gwynedd LL57 2UW, UK
2. Institute of Biodiversity, Animal Health & Comparative Medicine, University of Glasgow, Glasgow G12 8QQ, UK
3. Institute of Aquaculture, University of Stirling, Stirling FK9 4LA, UK
4. Ecosystems and Environment Research Centre, University of Salford, Salford M5 4WT, UK
5. School of Biological and Environmental Sciences, Liverpool John Moores University, Liverpool L3 5UG, UK