1. Department of Integrative Biology, University of California, Berkeley, CA 94720, USA
2. Mathematical Institute, University of Oxford, Oxford OX2 6GG, UK
3. College of Natural Resources, University of California, Berkeley, CA 94720, USA
4. College of Letters and Sciences, University of California, Berkeley, CA 94720, USA
5. Department of Science and Technology Studies, Division of Natural Science, York University, Toronto, Ontario, Canada M3J 1P3
6. Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter, Penryn Campus, Penryn TR10 9FE, UK