1. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, People's Republic of China
2. University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China
3. Jiangxi Province Key Laboratory of Watershed Ecosystem Change and Biodiversity, Institute of Life Science and School of Life Sciences, Nanchang University, Nanchang 330031, Jiangxi, People's Republic of China
4. Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany, Guangxi Zhuangzu Autonomous Region and the Chinese Academy of Sciences, Guilin 541006, Guangxi, People's Republic of China
5. Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d'Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, Paris 75005, France
6. Missouri Botanical Garden, 4344 Shaw Blvd, St Louis, MO 63166-0299, USA
7. Central Siberian Botanical Garden, Russian Academy of Sciences, Zolotodolinskaya Street 101, Novosibirsk 630090, Russia