1. Institute of Evolutionary Sciences of Montpellier (ISEM), UMR 5554, Université de Montpellier, CNRS, IRD, Montpellier, France
2. Department of Anthropology, Stony Brook University, Stony Brook, USA
3. Department of Anthropology, University College London, London, UK
4. Institute of Zoology, Zoological Society of London, London, UK