1. School of Ocean Sciences, Bangor University, Menai Bridge LL59 5AB, UK
2. Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth SY23 3DA, UK
3. Earth Institute and School of Biology and Environmental Science, University College Dublin, Dublin, Ireland
4. MaREI, the SFI Research Centre for Energy, Climate and Marine, Environmental Research Institute, University College Cork, Ringaskiddy, Ireland