1. Centre for Ecology and Conservation, University of Exeter, Cornwall Campus, Penryn, Cornwall TR10 9FE, UK
2. Swiss Ornithological Institute, Seerose 1, Sempach, 6204, Switzerland
3. Environment and Sustainability Institute, University of Exeter, Cornwall Campus, Penryn, Cornwall TR10 9FE, UK
4. Department of Entomology, Nanjing Agricultural University, Nanjing, 210095, People's Republic of China
5. Natural Resources Institute, University of Greenwich, Chatham, Kent SE10 9LS, UK
6. Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, UK