1. Department of Ecology, Radboud Institute for Biological and Environmental Sciences, Faculty of Science, Radboud University Nijmegen, 6525 AJ Nijmegen, The Netherlands
2. School of Biological Sciences, Faculty of Environmental and Life Sciences, University of Southampton, Southampton, SO17 1BJ, UK
3. Department of Ecosystem and Landscape Dynamics, Institute of Biodiversity and Ecosystem Dynamics (IBED-ELD), University of Amsterdam, P.O. Box 94240, 1090 GE Amsterdam, The Netherlands
4. Länsstyrelsen i Jönköpings län, Store Mosse Nationalpark, 335 74 Hillerstorp, Sweden