1. Department of Oceanography, School of Ocean and Earth Science and Technology (SOEST), University of Hawaiʻi at Mānoa, Honolulu, HI 96822, USA
2. Daniel K. Inouye Center for Microbial Oceanography: Research and Education, School of Ocean and Earth Science and Technology (SOEST), University of Hawaiʻi at Mānoa, Honolulu, HI 96822, USA
3. School of Oceanography, Shanghai Jiao Tong University, 1954 Huashan Rd, Shanghai Shi, Xuhui Qu 200240, China