1. National Socio-Environmental Synthesis Center SESYNC, 1 Park Place, Suite 300, Annapolis, MD 21401, USA
2. Department of Veterinary Population Medicine, University of Minnesota, St Paul, MN 55126, USA
3. Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Ashworth Labs, Charlotte Auerbach Road, Edinburgh EH9 3JT, UK