1. Department of Animal Ecology, Netherlands Institute of Ecology (NIOO-KNAW), 6708 PB, Wageningen, The Netherlands, The Netherlands
2. Behavioural Ecology Group, Wageningen University and Research, 6708 WD, Wageningen, The Netherlands
3. School of Biological Earth and Environmental Sciences, University College Cork, Cork, T23 N73K4, Ireland
4. Environmental Research Institute, University College Cork, Cork, T23 XE10, Ireland