1. Guangxi Key Laboratory for Forest Ecology and Conservation, College of Forestry, Guangxi University, Nanning 530004, People's Republic of China
2. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Institute of Biodiversity Science, Center of Evolutionary Biology, School of Life Sciences, Fudan University, Shanghai 200438, People's Republic of China
3. School of Life Science and Institute of Wetland Ecology, Nanjing University, Nanjing 210000, People's Republic of China
4. Biology Department, Edge Hill University, Ormskirk, Lancashire L39 4QP, UK
5. Key Laboratory of Environment Change and Resource Use, Beibu Gulf, Nanning Normal University, Nanning, Guangxi, People's Republic of China
6. Department of Biological Sciences, University of Cincinnati, Cincinnati OH 45221, USA