1. Montana Fish, Wildlife & Parks, 490 N. Meridian Rd. Kalispell, MT 59901, USA
2. Montana Fish, Wildlife & Parks, University of Montana, Fish Conservation Genetics Lab, Missoula, MT 59812, USA
3. U.S. Geological Survey, Northern Rocky Mountain Science Center, 38 Mather Dr., West Glacier, MT 59936, USA
4. Idaho Department of Fish and Game, 2885 W. Kathleen Ave., Coeur d'Alene, ID 83815, USA