1. Organismal and Evolutionary Biology, Faculty of Biological and Environmental Sciences, University of Helsinki, 00014, Finland
2. Tvärminne Zoological Station, University of Helsinki, J.A. Palménintie 260, 10900 Hanko, Finland
3. Faculty of Science, Ecology and Genetics Research Unit, University of Oulu, 90014, Finland
4. Laboratoire d'Ethologie Expérimentale et Comparée, UR 4443, Université Sorbonne Paris Nord, Villetaneuse, France
5. Institut Universitaire de France (IUF), Paris, France