1. Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, UK
2. Department of Mathematical Sciences, Loughborough University, Leicestershire LE11 3TU, UK
3. Graduate School of Public Health, Teikyo University, Tokyo 113-0033, Japan
4. Department of Data Science, Kitasato University, Kanagawa 252-0373, Japan
5. Centre for Biodiversity and Environmental Research, Department of Genetics, Evolution and Environment, University College London, Gower Street, WC1E 6BT London, UK