1. Department of Marine Ecology, Netherlands Institute for Sea Research (NIOZ), POBox 59, 1790 ABDen Burg, Texel, The Netherlands
2. Centre for Ecological and Evolutionary Studies (CEES), University of Groningen, POBox 14, 9750 AA Haren, The Netherlands
3. Department of Animal Sciences, University of Illinois, 1207 West Gregory Drive, Urbana, IL 61801, USA
4. Institute of Evolutionary and Ecological Sciences (IEEW), Leiden University, POBox 9516, 2300 RA Leiden, The Netherlands
5. Department of Physical Oceanography, Netherlands Institute for Sea Research (NIOZ), POBox 59, 1790 ABDen Burg, Texel, The Netherlands