1. Dipartimento di Etologia, Ecologia, Evoluzione, University of Pisa, Via A.Volta 6, I–56126 Pisa, Italy
2. School of Biological Sciences, University of WalesöSwansea, Singleton Park, Swansea SA2 8PP, UK
3. Centro di Studio per la Faunistica ed EcologiaTropicali del CNR, Via Romana 17, I–50125 Firenze, Italy
4. James Rennel Division, Southampton Oceanography Centre, Empress Dock, Southampton SO14 3ZH, UK