1. Australian School of Environmental Studies, Griffith University, Nathan, Queensland 4111, Australia
2. Netherlands Institute for Sea Research (NIOZ), PO Box 59, 1790 AB Den Burg, Texel, The Netherlands
3. Centre for Ecological and Evolutionary Studies, Zoological Laboratory, University of Groningen, PO Box 14, 9750 AA Haren, The Netherlands
4. Department of Biology, East China Normal University, Shanghai 200062, China