1. Department of Biology, University of Oulu, Post Office Box 3000, FIN–90014 Oulu, Finland
2. Institute of Zoology, Johannes Gutenberg University, Saarstrasse 21, D–55099 Mainz, Germany
3. Laboratory of Biodiversity Science, School of Agricultural and Life Sciences, University of Tokyo, Tokyo 113–8657, Japan
4. Institute of Biology and Soil Science, Far East Branch, RAS, 690022 Vladivostok>, Russia