1. Cardiff School of Biosciences, Cardiff University, Main Building, Park Place, Cardiff CF10 3TL, UK
2. School of Biological Sciences, Royal Holloway University of London, Egham, Surrey TW20 0EX, UK
3. Max–Planck–Institut für Verhaltensphysiologie, D–82319 Seewiesen, Germany
4. School of Biological Sciences, University of Bristol, Woodland Road, Bristol BS8 1UG, UK