1. The Marine Biological Association of the United Kingdom, The Laboratory, Citadel Hill, Plymouth PL1 2PB, UK
2. Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, PO Box 94766, 1090 GT Amsterdam, The Netherlands
3. Pisces Conservation Ltd, IRC House, The Square, Lymington, Hampshire SO41 8GN, UK
4. Biodiversity and Ecology Division, School of Biological Sciences, University of Southampton, Southampton SO16 7PX, UK