1. Department of Biology, Centre for Ecological and Evolutionary Synthesis, University of Oslo0316 Oslo, Norway
2. Department of Biological Sciences, University of ArkansasFayetteville, AR 72701, USA
3. Department of Biology, Microbial Evolution Research Group, University of Oslo0316 Oslo, Norway
4. Department of Zoology, University of OxfordSouth Parks Road, Oxford OX1 3PS, UK
5. Department of Zoology and Animal Biology, Molecular Systematics Group, University of Geneva1224 Chěne-Bougeries, Switzerland