1. Molecular and Evolutionary Ecology Group, Department of Biological Sciences, University of Hull, Hull HU6 7RX, UK
2. Institute of Evolutionary and Ecological Sciences, University of Leiden, PO Box 9516, 2300 RA Leiden, The Netherlands
3. Department of Zoology, University of Florida, Gainesville, FL 32611, USA
4. Department of Zoology, Makerere University, Kampala, Uganda
5. South African Institute for Aquatic Biodiversity, Somerset Street, Private Bag 1015, Grahamstown 6140, South Africa