1. Department of Biological and Environmental Science, University of Jyväauml;skyläauml;, PO Box 35, FIN–40014 Jyväauml;skyläauml;, Finland
2. School of Botany and Zoology, Australian National University, Canberra, ACT 0200, Australia ()
3. School of Biological, Earth and Environmental Science, The University of New South Wales, Sydney, NSW 2052, Australia ()
4. 116 Lynn Road, Ely, Cambridgeshire CB6 1DE, UK ()