1. Conservation Biology Group, Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
2. Zoological Museum, University of Copenhagen, Universitetsparken 15, DK–2100, Copenhagen Ø, Denmark
3. Biogeography and Conservation Laboratory, The Natural History Museum, Cromwell Road, London SW7 5BD, UK
4. Center for Applied Biodiversity Science, Conservation International, 1919 M Street NW, Suite 600, Washington, DC 20036, USA
5. 5Conservation Science Programme, World Wildlife Fund, 1250 24th Street NW, Washington, DC 20037-1193, USA
6. Llanunwas, Upper Solva, near Haverfordwest, Pembrokeshire SA62 6UJ, UK