1. Laboratoire de Biologie Animale, UMR 5555 du CNRS, Centre de Biologie et d'Ecologie Tropicale et Méditerranéenne, Universitéde Perpignan, 66860 Perpignan Cedex, France
2. School of Environmental Sciences and Development, Potchefstroom University, Private Bag X6001, Potchefstroom 2520, South Africa
3. Department of Microbiology and Parasitology, School of Molecular and Microbial Sciences, The University of Queensland, Brisbane, Queensland 4072, Australia