1. Institute of Evolutionary Biology, University of EdinburghKing's Buildings, West mains Road, Edinburgh EH9 3JT, UK
2. Macroecology and Conservation Unit, CEA—University of ÉvoraLargo dos Colegiais 2, 7002-554 Évora, Portugal
3. Department of Animal and Plant Sciences, University of SheffieldWestern Bank, Sheffield S10 2TN, UK
4. Population Biology, Department of Ecology and Evolution, Evolutionary Biology Centre, Uppsala UniversityNorbyvägen 18 D, 75236 Uppsala