1. Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, University of Oslo, PO Box 1050 Blindern, N-0316 Oslo, Norway
2. Department of Statistics and Actuarial Science, University of Iowa, Iowa City, IA 52242, USA
3. Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
4. Institute of Mathematics and Statistics, University of Kent at Canterbury, Canterbury CT2 7NF, UK
5. Department of Biological Sciences, Imperial College at Silwood Park, Ascot SL5 7PY, UK