1. Biodiversity and Macroecology Group, Department of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, UK
2. School of Biosciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, UK
3. National Institute for Public Health and the Environment (RIVM), Department for Environmental Information Systems (CIM), A. van Leeuwenhoeklaan 9, 3720 BA Bilthoven, The Netherlands