1. School of Biological Sciences, University of LiverpoolBiosciences Building, Crown Street, Liverpool L69 7ZB, UK
2. School of Biological Sciences, University of BristolWoodland Road, Bristol BS8 1UG, UK
3. Division of Environmental and Evolutionary Biology, Institute of Biomedical and Life Sciences, University of GlasgowGraham Kerr Building, Glasgow G12 8QQ, UK