1. Department of Mathematics, University College London, Gower Street, London WC1E 6BT, UK
2. Institute of Child Health, University College London, 30 Guilford Street, London WC1N 1EH, UK
3. CoMPLEX, Department of Biology, University College London, 4 Stephenson Way, London NW1 2HE, UK
4. The Galton Laboratory, Department of Biology, University College London, 4 Stephenson Way, London NW1 2HE, UK