1. Molecular Systematics Section, Jodrell LaboratoryRoyal Botanic Gardens, Kew, Richmond, Surrey TW9 3DS, UK
2. Department of Plant Biology, University of MinnesotaSaint Paul, MN 55108, USA
3. Department of Biological Sciences and NERC Centre for Population Biology, Imperial College LondonSilwood Park Campus, Ascot SL5 7PY, UK
4. Department of Botany, University of Dublin, Trinity CollegeDublin 2, UK
5. Department of Ecology and Evolutionary Biology, University of ArizonaTucson, AZ 85721, USA