1. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA 50011, USA
2. Bell Museum of Natural History, University of Minnesota, St. Paul, MN 55108, USA
3. Department of Fisheries, Wildlife, and Conservation Biology, University of Minnesota, St. Paul, MN 55108, USA
4. Department of Ecology and Evolution, Stony Brook University, Stony Brook, NY 11794-5245, USA