1. Institute of Cell, Animal and Population Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH9 3JT, UK
2. Agronomy and Range Science, University of California, 1 Shields Avenue, Davis, CA 95616, USA
3. Department of Integrative Biology, 3060 Valley Life Sciences Building No. 3140, University of California, Berkeley, CA 94720-3140, USA