1. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA
2. Conservation Genetics Group, Institute of Zoology, Regent's Park, London NW1 4RY, UK
3. Centre for Applied Conservation Biology, Forest Sciences, 2424 Main Mall, University of British Columbia, Vancouver, B.C., Canada, V6T1Z4
4. Cornell Laboratory of Ornithology, 159 Sapsucker Woods Road, Ithaca, NY 14850, USA
5. Centre for Biodiversity Research and Department of Zoology, University of British Columbia, Vancouver, B.C., Canada, V6T 1Z4